Notes on Uintacrinus socialis Grinnell
S. W. Williston. 1894.
Kansas University Quarterly,
Copyright ?2004-2010 by Mike Everhart
Page created 07/25/2004; Updated 12/22/2010
LEFT: Detail from a specimen of Uintacrinus socialis in the University of Kansas Museum of Natural History - Discovered by Prof. E. E. Slosson in 1891 (see below).
The only echinoderms known from the chalk are an unusual form of crinoid called Uintacrinus. Related to starfish and sea urchins, these animals were apparently colonial and free-living (not attached to the sea bottom). Uintacrinus socialis was first found in the Uinta Mountains of Utah by Marsh's party in 1870, but was mentioned only as a "new and very interesting crinoid, allied apparently to the Marsupites of the English Chalk" (Marsh, 1871, p. 195). Marsh's material from Utah was apparently so fragmentary that the species was actually described several years later largely from specimens found in the Smoky Hill Chalk of Kansas (Grinnell, 1876; Meek, 1876). In the chalk, Uintacrinus appears to have been somewhat colonial because its remains indicate many individuals died at the same time and settled to the sea bottom where they were preserved as thin layers of limestone. Miller, et al, (1957) reported that the slabs are usually between a quarter inch and one inch in thickness. Williston (1894) noted that he found a number of specimens in 1875 while collecting in the chalk with B. F. Mudge, and indicated that some of these served as the types for the genus and species described by Grinnell. One of Williston's students, B. H. Hill, wrote a description of the construction of the Late Cretaceous crinoid, Uintacrinus socialis, in 1894.
Notes on Uintacrinus socialis Grinnell.
While a member of the late Professor Mudge's party in western Kansas, during the summer of 1875, the writer was fortunate in finding in the Niobrara chalk a number of specimens of a crinoid which were notable from their very rarity. During that same season, whether before or after I do not now remember, other specimens of the same kind were discovered by Prof. Mudge and Mr. Geo. Cooper, all of which, as well as those found by myself, had been more or less exposed and weathered. A very few of these found their way into different collections, and among them were those which serve as the types of genus and species.* An imperfect specimen of this genus had been previously discovered by Marsh in the Uinta Mountains, but so incomplete that its affinities could not be decisively made out. ("In a stratum of yellow calcareous shale which overlies the coal series conformably, a thin layer was found full of Ostrea congesta Conrad, a typical Cretaceous fossil; and, just above, a new and very interesting crinoid, allied apparently to the Marsupites of the English chalk." Marsh, Amer. Journ. Sci., "March, 1871.) Because of this previous discovery, the generic name was chosen, but there is no proof that the species, at least, are identical.
During the season of 1891, Prof. E. E. Slosson, while a member of the Kansas University expedition in western Kansas, was so fortunate as to discover a most remarkable colony of this crinoid, by far the best yet known, in the vicinity of Elkader, on the Smoky Hill river. While all the colonies hitherto discovered have been exposed and more or less weathered, the present one was found in position, covered by the soft blue shale. The animals had lived so closely together that their very long arms had become inextricably entangled, and, by consolidation, had formed a dense calcareous plate, about one-third of an inch in thickness in the middle of the plate, but thinning out at the margin. About one-half of the thin slab as thus formed had been washed away; the remainder, as now restored in the University Museum, measures about six feet by three or four, and has, upon its under side, nearly one hundred of the crinoids, the greater part of which are perfectly preserved. The calyces all lie flattened out, showing, in some cases, the basal plates, but, as might be expected, never the upper or ventral
*Grinnell, Amer. Journ. Sci. xii, 81, July, 1876.
(19) KAN. UNIV. QUAR., VOL. II, NO. 1, JULY,1894.
20 KANSAS UNIVERSITY QUARTERLY.
portions. The interlacing of the arms prevents the tracing of any to the extremity. A photograph of a portion of the slab will be given in a future number of the QUARTERLY; for the present, the following description and figure, by Mr. B. H. Hill, student in paleontology in the University, will be of interest.
The horizon of Uintacrinus in Kansas seems to be confined to near the middle of the Niobrara. All the specimens hitherto discovered, of which I have any knowledge, have come from the vicinity of Elkader on the Smoky Hill, though in all probability, they will be found on both the Solomon and Saline.
S. W. WILLISTON.
In life, Uintacrinus socialis was evidently subglobose in shape, and about two inches in diameter. In place of the sub-basal plates of the stemmed crinoids, there is a small, five-sided, centro-dorsal plate, around which are grouped five pentagonal basals, the two
|longest sides of
which meet in a superior angle. The radials are fifteen in number, arranged in series of
three. The first radial is the broadest, broader than high, heptagonal in shape, the third
pentagonal. The two superior facets of the third radials give support to two series of
secondary radials, the proximal three of which are its supports. The arm plates are thin
and round, and radiate in structure. The arms themselves are ten in number.
NOTES ON UINTACRINUS SOCIALIS GRINNELL 21
In the colony there is one arm which I have traced for seventeen inches, and a comparison of its last distinguishable plates with the terminal ones of the other arms makes it evident that the arms must have been over two feet in length, as stated by Grinnell. They have well-developed pinnul?beginning at the base of the arm, where they are very large. They arise one from each plate, but alternating on the two sides of the ambulacral groove. Grinnell described what he believed to be interbrachial and interradial arms, but it seems certain that he was in error in this respect, having mistaken the pinnul?for these arms. The interradials are usually seven in number. The first is hexagonal and lies between the first and second radials of two series. The other six are also hexagonal, and in life may have been arranged in pairs, but are preserved in the specimens in irregular groups of four. The interbrachials are two in number, heptagonal in shape, and lie, respectively, between the first and second, and the second and third secondary radials of the two arms.
From the shape of the crinoid, its globose form and long, heavy arms, one would hardly expect to find any of the ventral plates exposed, and such is the case. Nor has it been possible to expose them by dissecting away the plates.
B. H. HILL.
|LEFT: A detail from the beautiful exhibit specimen of
socialis at the Sternberg Museum. This specimen was found by George F. Sternberg in
Logan County. See other close-ups of individual crinoids in this specimen: HERE, HERE and HERE.
RIGHT: Figures from Meek (1876) of two specimens collected by B.F. Mudge in the Smoky Hill Chalk of Western Kansas.
"Their bodies were about the shape of half an egg, with an opening in the center, and ten arms radiating from the margin. These arms were three feet long, with feathered edges. Over the mouth, too, were smaller arms used to comb off into the mouth the tiny animal life of the sea, that was strained through, and caught in the meshes of the feathered arms. My boys found hundreds of these crinoids in the Chalk on Beaver Creek, Kansas, called Uintacrinus socialis. We enriched many Museums with them."
Charles H. Sternberg's "Hunting Dinosaurs on the Red Deer River, Alberta, Canada" (1917, p. 156).
Beecher, C. E. 1900. On a large slab of Uintacrinus from Kansas. American Journal Science 18:23-24, 2 pl. (Specimen at Yale University, collected by H. T. Martin)
Grinnell, G. B. 1876. On a new crinoid from the Cretaceous formation of the west. American Journal of Science ser. 3, 12(3):81-83.
Grinnell, G. B. 1894. Notes on Uintacrinus socialis Grinnell. Kansas University Quarterly 3(1):19-21.
Marsh, O. C. 1871. On the geology of the eastern Uintah Mountains. American Journal of Science, Series 3, 1(3):191-198. (Uintacrinus)
Martin, H. T. 1907. Some new features of Uintacrinus. University Kansas Science Bulletin 4(6):193-196, pl. IX-X.Meek, F.B. 1876. Note on the new genus Uintacrinus, Grinnell. Bulletin
Meyer, D. L. and C. V. Milsom. 2001. Microbial sealing in the biostratinomy of Uintacrinus Lagerst鋞ten in the upper Cretaceous of Kansas and Colorado, USA. Palaios 16:535-546.
Miller, H. W. Jr., G. F. Sternberg and M. V. Walker. 1957. Uintacrinus localities in the Niobrara Formation of Kansas. Kansas Academy of Science, Transactions 60(2):163-166, 2 fig.
Mudge, B. F. 1877. Annual Report of the Committee on Geology, for the year ending November 1, 1876. Kansas Academy of Science, Transactions, Ninth Annual Meeting, pp. 4-5. (discovery of Uintacrinus socialis in Kansas, Pteranodon, sharks and birds.)
Springer, F. 1889. Notice of a new discovery concerning Uintacrinus. American Geologist 24(2):92.
Sternberg, C. H. 1917. Hunting Dinosaurs on the Red Deer River, Alberta, Canada. Published by the author.
Williston, S. W. 1894. Notes on Uintacrinus socialis Grinnell. Kansas University Quarterly, 3(1):19-21 (includes note by B. H. Hill).